第 29 节
作者:无组织      更新:2022-04-21 11:08      字数:9322
  tells us; on the first page of his preface; that he has no intention
  of dealing even in outline with the vast subject of evolution in
  general; and has only tried to give such an account of the theory of
  natural selection as may facilitate a clear conception of Darwin's
  work。  How far he has succeeded is a point on which opinion will
  probably be divided。  Those who find Mr。 Darwin's works clear will
  also find no difficulty in understanding Mr。 Wallace; those; on the
  other hand; who find Mr。 Darwin puzzling are little likely to be
  less puzzled by Mr。 Wallace。  He continues:…
  〃The objections now made to Darwin's theory apply solely to the
  particular means by which the change of species has been brought
  about; not to the fact of that change。〃
  But 〃Darwin's theory〃as Mr。 Wallace has elsewhere proved that he
  understandshas no reference 〃to the fact of that change〃that is
  to say; to the fact that species have been modified in course of
  descent from other species。  This is no more Mr。 Darwin's theory
  than it is the reader's or my own。  Darwin's theory is concerned
  only with 〃the particular means by which the change of species has
  been brought about〃; his contention being that this is mainly due to
  the natural survival of those individuals that have happened by some
  accident to be born most favourably adapted to their surroundings;
  or; in other words; through accumulation in the common course of
  nature of the more lucky variations that chance occasionally
  purveys。  Mr。 Wallace's words; then; in reality amount to this; that
  the objections now made to Darwin's theory apply solely to Darwin's
  theory; which is all very well as far as it goes; but might have
  been more easily apprehended if he had simply said; 〃There are
  several objections now made to Mr。 Darwin's theory。〃
  It must be remembered that the passage quoted above occurs on the
  first page of a preface dated March 1889; when the writer had
  completed his task; and was most fully conversant with his subject。
  Nevertheless; it seems indisputable either that he is still
  confusing evolution with Mr。 Darwin's theory; or that he does not
  know when his sentences have point and when they have none。
  I should perhaps explain to some readers that Mr。 Darwin did not
  modify the main theory put forward; first by Buffon; to whom it
  indisputably belongs; and adopted from him by Erasmus Darwin;
  Lamarck; and many other writers in the latter half of the last
  century and the earlier years of the present。  The early
  evolutionists maintained that all existing forms of animal and
  vegetable life; including man; were derived in course of descent
  with modification from forms resembling the lowest now known。
  Mr。 Darwin went as far as this; and farther no one can go。  The
  point at issue between him and his predecessors involves neither the
  main fact of evolution; nor yet the geometrical ratio of increase;
  and the struggle for existence consequent thereon。  Messrs。 Darwin
  and Wallace have each thrown invaluable light upon these last two
  points; but Buffon; as early as 1756; had made them the keystone of
  his system。  〃The movement of nature;〃 he then wrote; 〃turns on two
  immovable pivots:  one; the illimitable fecundity which she has
  given to all species:  the other; the innumerable difficulties which
  reduce the results of that fecundity。〃  Erasmus Darwin and Lamarck
  followed in the same sense。  They thus admit the survival of the
  fittest as fully as Mr。 Darwin himself; though they do not make use
  of this particular expression。  The dispute turns not upon natural
  selection; which is common to all writers on evolution; but upon the
  nature and causes of the variations that are supposed to be selected
  from and thus accumulated。  Are these mainly attributable to the
  inherited effects of use and disuse; supplemented by occasional
  sports and happy accidents?  Or are they mainly due to sports and
  happy accidents; supplemented by occasional inherited effects of use
  and disuse?
  The Lamarckian system has all along been maintained by Mr。 Herbert
  Spencer; who; in his 〃Principles of Biology;〃 published in 1865;
  showed how impossible it was that accidental variations should
  accumulate at all。  I am not sure how far Mr。 Spencer would consent
  to being called a Lamarckian pure and simple; nor yet how far it is
  strictly accurate to call him one; nevertheless; I can see no
  important difference in the main positions taken by him and by
  Lamarck。
  The question at issue between the Lamarckians; supported by Mr。
  Spencer and a growing band of those who have risen in rebellion
  against the Charles…Darwinian system on the one hand; and Messrs。
  Darwin and Wallace with the greater number of our more prominent
  biologists on the other; involves the very existence of evolution as
  a workable theory。  For it is plain that what Nature can be supposed
  able to do by way of choice must depend on the supply of the
  variations from which she is supposed to choose。  She cannot take
  what is not offered to her; and so again she cannot be supposed able
  to accumulate unless what is gained in one direction in one
  generation; or series of generations; is little likely to be lost in
  those that presently succeed。  Now variations ascribed mainly to use
  and disuse can be supposed capable of being accumulated; for use and
  disuse are fairly constant for long periods among the individuals of
  the same species; and often over large areas; moreover; conditions
  of existence involving changes of habit; and thus of organisation;
  come for the most part gradually; so that time is given during which
  the organism can endeavour to adapt itself in the requisite
  respects; instead of being shocked out of existence by too sudden
  change。  Variations; on the other hand; that are ascribed to mere
  chance cannot be supposed as likely to be accumulated; for chance is
  notoriously inconstant; and would not purvey the variations in
  sufficiently unbroken succession; or in a sufficient number of
  individuals; modified similarly in all the necessary correlations at
  the same time and place to admit of their being accumulated。  It is
  vital therefore to the theory of evolution; as was early pointed out
  by the late Professor Fleeming Jenkin and by Mr。 Herbert Spencer;
  that variations should be supposed to have a definite and persistent
  principle underlying them; which shall tend to engender similar and
  simultaneous modification; however small; in the vast majority of
  individuals composing any species。  The existence of such a
  principle and its permanence is the only thing that can be supposed
  capable of acting as rudder and compass to the accumulation of
  variations; and of making it hold steadily on one course for each
  species; till eventually many havens; far remote from one another;
  are safely reached。
  It is obvious that the having fatally impaired the theory of his
  predecessors could not warrant Mr。 Darwin in claiming; as he most
  fatuously did; the theory of evolution。  That he is still generally
  believed to have been the originator of this theory is due to the
  fact that he claimed it; and that a powerful literary backing at
  once came forward to support him。  It seems at first sight
  improbable that those who too zealously urged his claims were
  unaware that so much had been written on the subject; but when we
  find even Mr。 Wallace himself as profoundly ignorant on this subject
  as he still either is; or affects to be; there is no limit
  assignable to the ignorance or affected ignorance of the kind of
  biologists who would write reviews in leading journals thirty years
  ago。  Mr。 Wallace writes:…
  〃A few great naturalists; struck by the very slight difference
  between many of these species; and the numerous links that exist
  between the most different forms of animals and plants; and also
  observing that a great many species do vary considerably in their
  forms; colours and habits; conceived the idea that they might be all
  produced one from the other。  The most eminent of these writers was
  a great French naturalist; Lamarck; who published an elaborate work;
  the Philosophie Zoologique; in which he endeavoured to prove that
  all animals whatever are descended from other species of animals。
  He attributed the change of species chiefly to the effect of changes
  in the conditions of lifesuch as climate; food; &c。; and
  especially to the desires and efforts of the animals themselves to
  improve their condition; leading to a modification of form or size
  in certain parts; owing to the well…known physiological law that all
  organs are strengthened by constant use; while they are weakened or
  even completely lost by disuse 。 。 。
  〃The only other important work dealing with the question was the
  celebrated 'Vestiges of Creation;' published anonymously; but now
  acknowledged to have been written by the late Robert Chambers。〃
  None are so blind as those who will not see; and it would be waste
  of time to argue with the invincible ignorance of one who thinks
  Lamarck and Buffon conceived that all species were produced from one
  another; more especially as I have already dealt at some length with
  the early evolutionists in my work; 〃Evolution; Old and New;〃 first
  published ten years ago;