第 29 节
作者:猜火车      更新:2021-02-24 23:33      字数:9322
  the oxide。  Acetic acid produced hardly any effect on his sand; and
  even hydrochloric; nitric and sulphuric acids; diluted as in the
  Pharmacopoeia; produced less effect than did the acids in the
  intestines of the worms。
  Mr。 A。 A。 Julien has lately collected all the extant information
  about the acids generated in humus; which; according to some
  chemists; amount to more than a dozen different kinds。  These
  acids; as well as their acid salts (i。e。; in combination with
  potash; soda; and ammonia); act energetically on carbonate of lime
  and on the oxides of iron。  It is also known that some of these
  acids; which were called long ago by Thenard azohumic; are enabled
  to dissolve colloid silica in proportion to the nitrogen which they
  contain。 {64}  In the formation of these latter acids worms
  probably afford some aid; for Dr。 H。 Johnson informs me that by
  Nessler's test he found 0。018 per cent。 of ammonia in their
  castings。
  It may be here added that I have recently been informed by Dr。
  Gilbert 〃that several square yards on his lawn were swept clean;
  and after two or three weeks all the worm…castings on the space
  were collected and dried。  These were found to contain 0。35 of
  nitrogen。  This is from two to three times as much as we find in
  our ordinary arable surface…soil; more than in our ordinary pasture
  surface…soil; but less than in rich kitchen…garden mould。
  Supposing a quantity of castings equal to 10 tons in the dry state
  were annually deposited on an acre; this would represent a manuring
  of 78 lbs。 of nitrogen per acre per annum; and this is very much
  more than the amount of nitrogen in the annual yield of hay per
  acre; if raised without any nitrogenous manure。  Obviously; so far
  as the nitrogen in the castings is derived from surface…growth or
  from surface…soil; it is not a gain to the latter; but so far as it
  is derived from below; it is a gain。〃
  The several humus…acids; which appear; as we have just seen; to be
  generated within the bodies of worms during the digestive process;
  and their acid salts; play a highly important part; according to
  the recent observations of Mr。 Julien; in the disintegration of
  various kinds of rocks。  It has long been known that the carbonic
  acid; and no doubt nitric and nitrous acids; which are present in
  rain…water; act in like manner。  There is; also; a great excess of
  carbonic acid in all soils; especially in rich soils; and this is
  dissolved by the water in the ground。  The living roots of plants;
  moreover; as Sachs and others have shown; quickly corrode and leave
  their impressions on polished slabs of marble; dolomite and
  phosphate of lime。  They will attack even basalt and sandstone。
  {65}  But we are not here concerned with agencies which are wholly
  independent of the action of worms。
  The combination of any acid with a base is much facilitated by
  agitation; as fresh surfaces are thus continually brought into
  contact。  This will be thoroughly effected with the particles of
  stone and earth in the intestines of worms; during the digestive
  process; and it should be remembered that the entire mass of the
  mould over every field; passes; in the course of a few years;
  through their alimentary canals。  Moreover as the old burrows
  slowly collapse; and as fresh castings are continually brought to
  the surface; the whole superficial layer of mould slowly revolves
  or circulates; and the friction of the particles one with another
  will rub off the finest films of disintegrated matter as soon as
  they are formed。  Through these several means; minute fragments of
  rocks of many kinds and mere particles in the soil will be
  continually exposed to chemical decomposition; and thus the amount
  of soil will tend to increase。
  As worms line their burrows with their castings; and as the burrows
  penetrate to a depth of 5 or 6; or even more feet; some small
  amount of the humus…acids will be carried far down; and will there
  act on the underlying rocks and fragments of rock。  Thus the
  thickness of the soil; if none be removed from the surface; will
  steadily though slowly tend to increase; but the accumulation will
  after a time delay the disintegration of the underlying rocks and
  of the more deeply seated particles。  For the humus…acids which are
  generated chiefly in the upper layer of vegetable mould; are
  extremely unstable compounds; and are liable to decomposition
  before they reach any considerable depth。 {66}  A thick bed of
  overlying soil will also check the downward extension of great
  fluctuations of temperature; and in cold countries will check the
  powerful action of frost。  The free access of air will likewise be
  excluded。  From these several causes disintegration would be almost
  arrested; if the overlying mould were to increase much in
  thickness; owing to none or little being removed from the surface。
  {67}  In my own immediate neighbourhood we have a curious proof how
  effectually a few feet of clay checks some change which goes on in
  flints; lying freely exposed; for the large ones which have lain
  for some time on the surface of ploughed fields cannot be used for
  building; they will not cleave properly; and are said by the
  workmen to be rotten。 {68}  It is therefore necessary to obtain
  flints for building purposes from the bed of red clay overlying the
  chalk (the residue of its dissolution by rain…water) or from the
  chalk itself。
  Not only do worms aid directly in the chemical disintegration of
  rocks; but there is good reason to believe that they likewise act
  in a direct and mechanical manner on the smaller particles。  All
  the species which swallow earth are furnished with gizzards; and
  these are lined with so thick a chitinous membrane; that Perrier
  speaks of it; {69} as 〃une veritable armature。〃  The gizzard is
  surrounded by powerful transverse muscles; which; according to
  Claparede; are about ten times as thick as the longitudinal ones;
  and Perrier saw them contracting energetically。  Worms belonging to
  one genus; Digaster; have two distinct but quite similar gizzards;
  and in another genus; Moniligaster; the second gizzard consists of
  four pouches; one succeeding the other; so that it may almost be
  said to have five gizzards。 {70}  In the same manner as
  gallinaceous and struthious birds swallow stones to aid in the
  trituration of their food; so it appears to be with terricolous
  worms。  The gizzards of thirty…eight of our common worms were
  opened; and in twenty…five of them small stones or grains of sand;
  sometimes together with the hard calcareous concretions formed
  within the anterior calciferous glands; were found; and in two
  others concretions alone。  In the gizzards of the remaining worms
  there were no stones; but some of these were not real exceptions;
  as the gizzards were opened late in the autumn; when the worms had
  ceased to feed and their gizzards were quite empty。 {71}
  When worms make their burrows through earth abounding with little
  stones; no doubt many will be unavoidably swallowed; but it must
  not be supposed that this fact accounts for the frequency with
  which stones and sand are found in their gizzards。  For beads of
  glass and fragments of brick and of hard tiles were scattered over
  the surface of the earth; in pots in which worms were kept and had
  already made their burrows; and very many of these beads and
  fragments were picked up and swallowed by the worms; for they were
  found in their castings; intestines; and gizzards。  They even
  swallowed the coarse red dust; formed by the pounding of the tiles。
  Nor can it be supposed that they mistook the beads and fragments
  for food; for we have seen that their taste is delicate enough to
  distinguish between different kinds of leaves。  It is therefore
  manifest that they swallow hard objects; such as bits of stone;
  beads of glass and angular fragments of bricks or tiles for some
  special purpose; and it can hardly be doubted that this is to aid
  their gizzards in crushing and grinding the earth; which they so
  largely consume。  That such hard objects are not necessary for
  crushing leaves; may be inferred from the fact that certain
  species; which live in mud or water and feed on dead or living
  vegetable matter; but which do not swallow earth; are not provided
  with gizzards; {72} and therefore cannot have the power of
  utilising stones。
  During the grinding process; the particles of earth must be rubbed
  against one another; and between the stones and the tough lining
  membrane of the gizzard。  The softer particles will thus suffer
  some attrition; and will perhaps even be crushed。  This conclusion
  is supported by the appearance of freshly ejected castings; for
  these often reminded me of the appearance of paint which has just
  been ground by a workman between two flat stones。  Morren remarks
  that the intestinal canal is 〃impleta tenuissima terra; veluti in
  pulverem redacta。〃 {73}  Perrier also speaks of 〃l'etat de pate
  excessivement fine a laquelle est reduite la terre qu'ils
  rejettent;〃 &c。 {74}
  As the amount of trituration which the particles of earth undergo
  in the gizzards of worms pos