第 8 节
作者:猜火车      更新:2021-02-24 23:33      字数:9321
  burrows are not gnawed。  I have seen as many as nine leaves of the
  lime…tree drawn into the same burrow; and not nearly all of them
  had been gnawed; but such leaves may serve as a store for future
  consumption。  Where fallen leaves are abundant; many more are
  sometimes collected over the mouth of a burrow than can be used; so
  that a small pile of unused leaves is left like a roof over those
  which have been partly dragged in。
  A leaf in being dragged a little way into a cylindrical burrow is
  necessarily much folded or crumpled。  When another leaf is drawn
  in; this is done exteriorly to the first one; and so on with the
  succeeding leaves; and finally all become closely folded and
  pressed together。  Sometimes the worm enlarges the mouth of its
  burrow; or makes a fresh one close by; so as to draw in a still
  larger number of leaves。  They often or generally fill up the
  interstices between the drawn…in leaves with moist viscid earth
  ejected from their bodies; and thus the mouths of the burrows are
  securely plugged。  Hundreds of such plugged burrows may be seen in
  many places; especially during the autumnal and early winter
  months。  But; as will hereafter be shown; leaves are dragged into
  the burrows not only for plugging them up and for food; but for the
  sake of lining the upper part or mouth。
  When worms cannot obtain leaves; petioles; sticks; &c。; with which
  to plug up the mouths of their burrows; they often protect them by
  little heaps of stones; and such heaps of smooth rounded pebbles
  may frequently be seen on gravel…walks。  Here there can be no
  question about food。  A lady; who was interested in the habits of
  worms; removed the little heaps of stones from the mouths of
  several burrows and cleared the surface of the ground for some
  inches all round。  She went out on the following night with a
  lantern; and saw the worms with their tails fixed in their burrows;
  dragging the stones inwards by the aid of their mouths; no doubt by
  suction。  〃After two nights some of the holes had 8 or 9 small
  stones over them; after four nights one had about 30; and another
  34 stones。〃 {29}  One stonewhich had been dragged over the
  gravel…walk to the mouth of a burrow weighed two ounces; and this
  proves how strong worms are。  But they show greater strength in
  sometimes displacing stones in a well…trodden gravel…walk; that
  they do so; may be inferred from the cavities left by the displaced
  stones being exactly filled by those lying over the mouths of
  adjoining burrows; as I have myself observed。
  Work of this kind is usually performed during the night; but I have
  occasionally known objects to be drawn into the burrows during the
  day。  What advantage the worms derive from plugging up the mouths
  of their burrows with leaves; &c。; or from piling stones over them;
  is doubtful。  They do not act in this manner at the times when they
  eject much earth from their burrows; for their castings then serve
  to cover the mouths。  When gardeners wish to kill worms on a lawn;
  it is necessary first to brush or rake away the castings from the
  surface; in order that the lime…water may enter the burrows。 {30}
  It might be inferred from this fact that the mouths are plugged up
  with leaves; &c。; to prevent the entrance of water during heavy
  rain; but it may be urged against this view that a few; loose;
  well…rounded stones are ill…adapted to keep out water。  I have
  moreover seen many burrows in the perpendicularly cut turf…edgings
  to gravel…walks; into which water could hardly flow; as well
  plugged as burrows on a level surface。  It is not probable that the
  plugs or piles of stones serve to conceal the burrows from
  scolopendras; which; according to Hoffmeister; {31} are the
  bitterest enemies of worms; or from the larger species of Carabus
  and Staphylinus which attack them ferociously; for these animals
  are nocturnal; and the burrows are opened at night。  May not worms
  when the mouth of the burrow is protected be able to remain with
  safety with their heads close to it; which we know that they like
  to do; but which costs so many of them their lives?  Or may not the
  plugs check the free ingress of the lowest stratum of air; when
  chilled by radiation at night; from the surrounding ground and
  herbage?  I am inclined to believe in this latter view:  firstly;
  because when worms were kept in pots in a room with a fire; in
  which case cold air could not enter the burrows; they plugged them
  up in a slovenly manner; and secondarily; because they often coat
  the upper part of their burrows with leaves; apparently to prevent
  their bodies from coming into close contact with the cold damp
  earth。  Mr。 E。 Parfitt has suggested to me that the mouths of the
  burrows are closed in order that the air within them may be kept
  thoroughly damp; and this seems the most probable explanation of
  the habit。  But the plugging…up process may serve for all the above
  purposes。
  Whatever the motive may be; it appears that worms much dislike
  leaving the mouths of their burrows open。  Nevertheless they will
  reopen them at night; whether or not they can afterwards close
  them。  Numerous open burrows may be seen on recently…dug ground;
  for in this case the worms eject their castings in cavities left in
  the ground; or in the old burrows instead of piling them over the
  mouths of their burrows; and they cannot collect objects on the
  surface by which the mouths might be protected。  So again on a
  recently disinterred pavement of a Roman villa at Abinger
  (hereafter to be described) the worms pertinaciously opened their
  burrows almost every night; when these had been closed by being
  trampled on; although they were rarely able to find a few minute
  stones wherewith to protect them。
  Intelligence shown by worms in their manner of plugging up their
  burrows。If a man had to plug up a small cylindrical hole; with
  such objects as leaves; petioles or twigs; he would drag or push
  them in by their pointed ends; but if these objects were very thin
  relatively to the size of the hole; he would probably insert some
  by their thicker or broader ends。  The guide in his case would be
  intelligence。  It seemed therefore worth while to observe carefully
  how worms dragged leaves into their burrows; whether by their tips
  or bases or middle parts。  It seemed more especially desirable to
  do this in the case of plants not natives to our country; for
  although the habit of dragging leaves into their burrows is
  undoubtedly instinctive with worms; yet instinct could not tell
  them how to act in the case of leaves about which their progenitors
  knew nothing。  If; moreover; worms acted solely through instinct or
  an unvarying inherited impulse; they would draw all kinds of leaves
  into their burrows in the same manner。  If they have no such
  definite instinct; we might expect that chance would determine
  whether the tip; base or middle was seized。  If both these
  alternatives are excluded; intelligence alone is left; unless the
  worm in each case first tries many different methods; and follows
  that alone which proves possible or the most easy; but to act in
  this manner and to try different methods makes a near approach to
  intelligence。
  In the first place 227 withered leaves of various kinds; mostly of
  English plants; were pulled out of worm…burrows in several places。
  Of these; 181 had been drawn into the burrows by or near their
  tips; so that the foot…stalk projected nearly upright from the
  mouth of the burrow; 20 had been drawn in by their bases; and in
  this case the tips projected from the burrows; and 26 had been
  seized near the middle; so that these had been drawn in
  transversely and were much crumpled。  Therefore 80 per cent。
  (always using the nearest whole number) had been drawn in by the
  tip; 9 per cent。 by the base or foot…stalk; and 11 per cent。
  transversely or by the middle。  This alone is almost sufficient to
  show that chance does not determine the manner in which leaves are
  dragged into the burrows。
  Of the above 227 leaves; 70 consisted of the fallen leaves of the
  common lime…tree; which is almost certainly not a native of
  England。  These leaves are much acuminated towards the tip; and are
  very broad at the base with a well…developed foot…stalk。  They are
  thin and quite flexible when half…withered。  Of the 70; 79 per
  cent。 had been drawn in by or near the tip; 4 per cent。 by or near
  the base; and 17 per cent。 transversely or by the middle。  These
  proportions agree very closely; as far as the tip is concerned;
  with those before given。  But the percentage drawn in by the base
  is smaller; which may be attributed to the breadth of the basal
  part of the blade。  We here; also; see that the presence of a foot…
  stalk; which it might have been expected would have tempted the
  worms as a convenient handle; has little or no influence in
  determining the manner in which lime leaves are dragged into the
  burrows。  The considerable proportion; viz。; 17 per cent。; drawn in
  more or less transversely depends no doubt on the flexibility of
  these half…decayed leaves。  The fact of so many having been drawn