第 25 节
作者:
匆匆 更新:2024-04-07 11:54 字数:9322
complicated activities of man are comprehensible under three
categories。 Either they are immediately directed towards the
maintenance and development of the body; or they effect transitory
changes in the relative positions of parts of the body; or they
tend towards the continuance of the species。 Even those
manifestations of intellect; of feeling; and of will; which we
rightly name the higher faculties; are not excluded from this
classification; inasmuch as to every one but the subject of them;
they are known only as transitory changes in the relative positions
of parts of the body。 Speech; gesture; and every other form of
human action are; in the long run; resolvable into muscular
contraction; and muscular contraction is but a transitory change in
the relative positions of the parts of a muscle。 But the scheme
which is large enough to embrace the activities of the highest form
of life; covers all those of the lower creatures。 The lowest
plant; or animalcule; feeds; grows; and reproduces its kind。 In
addition; all animals manifest those transitory changes of form
which we class under irritability and contractility; and; it is
more than probable; that when the vegetable world is thoroughly
explored; we shall find all plants in possession of the same
powers; at one time or other of their existence。
I am not now alluding to such phaenomena; at once rare and
conspicuous; as those exhibited by the leaflets of the sensitive
plants; or the stamens of the barberry; but to much more widely
spread; and at the same time; more subtle and hidden;
manifestations of vegetable contractility。 You are doubtless aware
that the common nettle owes its stinging property to the
innumerable stiff and needle…like; though exquisitely delicate;
hairs which cover its surface。 Each stinging…needle tapers from a
broad base to a slender summit; which; though rounded at the end;
is of such microscopic fineness that it readily penetrates; and
breaks off in; the skin。 The whole hair consists of a very
delicate outer case of wood; closely applied to the inner surface
of which is a layer of semi…fluid matter; full of innumerable
granules of extreme minuteness。 This semi…fluid lining is
protoplasm; which thus constitutes a kind of bag; full of a limpid
liquid; and roughly corresponding in form with the interior of the
hair which it fills。 When viewed with a sufficiently high
magnifying power; the protoplasmic layer of the nettle hair is seen
to be in a condition of unceasing activity。 Local contractions of
the whole thickness of its substance pass slowly and gradually from
point to point; and give rise to the appearance of progressive
waves; just as the bending of successive stalks of corn by a breeze
produces the apparent billows of a cornfield。
But; in addition to these movements; and independently of them; the
granules are driven; in relatively rapid streams; through channels
in the protoplasm which seem to have a considerable amount of
persistence。 Most commonly; the currents in adjacent parts of the
protoplasm take similar directions; and; thus; there is a general
stream up one side of the hair and down the other。 But this does
not prevent the existence of partial currents which take different
routes; and sometimes trains of granules may be seen coursing
swiftly in opposite directions within a twenty…thousandth of an
inch of one another; while; occasionally; opposite streams come
into direct collision; and; after a longer or shorter struggle; one
predominates。 The cause of these currents seems to lie in
contractions of the protoplasm which bounds the channels in which
they flow; but which are so minute that the best microscopes show
only their effects; and not themselves。
The spectacle afforded by the wonderful energies prisoned within
the compass of the microscopic hair of a plant; which we commonly
regard as a merely passive organism; is not easily forgotten by one
who has watched its display; continued hour after hour; without
pause or sign of weakening。 The possible complexity of many other
organic forms; seemingly as simple as the protoplasm of the nettle;
dawns upon one; and the comparison of such a protoplasm to a body
with an internal circulation; which has been put forward by an
eminent physiologist; loses much of its startling character。
Currents similar to those of the hairs of the nettle have been
observed in a great multitude of very different plants; and weighty
authorities have suggested that they probably occur; in more or
less perfection; in all young vegetable cells。 If such be the
case; the wonderful noonday silence of a tropical forest is; after
all; due only to the dulness of our hearing; and could our ears
catch the murmur of these tiny Maelstroms; '96' as they whirl in the
innumerable myriads of living cells which constitute each tree; we
should be stunned; as with the roar of a great city。
Among the lower plants; it is the rule rather than the exception;
that contractility should be still more openly manifested at some
periods of their existence。 The protoplasm of Algae and Fungi
becomes; under many circumstances; partially; or completely; freed
from its woody case; and exhibits movements of its whole mass; or
is propelled by the contractility of one; or more; hair…like
prolongations of its body; which are called vibratile cilia。 And;
so far as the conditions of the manifestation of the phaenomena of
contractility have yet been studied; they are the same for the
plant as for the animal。 Heat and electric shocks influence both;
and in the same way; though it may be in different degrees。 It is
by no means my intention to suggest that there is no difference in
faculty between the lowest plant and the highest; or between plants
and animals。 But the difference between the powers of the lowest
plant; or animal; and those of the highest; is one of degree; not
of kind; and depends; as Milne…Edwards '97' long ago so well pointed
out; upon the extent to which the principle of the division of
labour is carried out in the living economy。 In the lowest
organism all parts are competent to perform all functions; and one
and the same portion of protoplasm may successfully take on the
function of feeding; moving; or reproducing apparatus。 In the
highest; on the contrary; a great number of parts combine to
perform each function; each part doing its allotted share of the
work with great accuracy and efficiency; but being useless for any
other purpose。
On the other hand; notwithstanding all the fundamental resemblances
which exist between the powers of the protoplasm in plants and in
animals; they present a striking difference (to which I shall
advert more at length presently); in the fact that plants can
manufacture fresh protoplasm out of mineral compounds; whereas
animals are obliged to procure it ready made; and hence; in the
long run; depend upon plants。 Upon what condition this difference
in the powers of the two great divisions of the world of life
depends; nothing is at present known。
With such qualifications as arises '98' out of the last…mentioned
fact; it may be truly said that the acts of all living things are
fundamentally one。 Is any such unity predicable of their forms?
Let us seek in easily verified facts for a reply to this question。
If a drop of blood be drawn by pricking one's finger; and viewed
with proper precautions; and under a sufficiently high microscopic
power; there will be seen; among the innumerable multitude of
little; circular; discoidal bodies; or corpuscles; which float in
it and give it its colour; a comparatively small number of
colourless corpuscles; of somewhat larger size and very irregular
shape。 If the drop of blood be kept at the temperature of the
body; these colourless corpuscles will be seen to exhibit a
marvellous activity; changing their forms with great rapidity;
drawing in and thrusting out prolongations of their substance; and
creeping about as if they were independent organisms。
The substance which is thus active is a mass of protoplasm; and its
activity differs in detail; rather than in principle; from that of
the protoplasm of the nettle。 Under sundry circumstances the
corpuscle dies and becomes distended into a round mass; in the
midst of which is seen a smaller spherical body; which existed; but
was more or less hidden; in the living corpuscle; and is called its
nucleus。 Corpuscles of essentially similar structure are to be
found in the skin; in the lining of the mouth; and scattered
through the whole framework of the body。 Nay; more; in the
earliest condition of the human organism; in that state in which it
has but just become distinguishable from the egg in which it
arises; it is nothing but an aggregation of such corpuscles; and
every organ of the body was; once; no more than such an
aggregation。
Thus a nucleated mass of protoplasm tur