第 25 节
作者:匆匆      更新:2024-04-07 11:54      字数:9322
  complicated activities of man are comprehensible under three
  categories。  Either they are immediately directed towards the
  maintenance and development of the body; or they effect transitory
  changes in the relative positions of parts of the body; or they
  tend towards the continuance of the species。  Even those
  manifestations of intellect; of feeling; and of will; which we
  rightly name the higher faculties; are not excluded from this
  classification; inasmuch as to every one but the subject of them;
  they are known only as transitory changes in the relative positions
  of parts of the body。  Speech; gesture; and every other form of
  human action are; in the long run; resolvable into muscular
  contraction; and muscular contraction is but a transitory change in
  the relative positions of the parts of a muscle。  But the scheme
  which is large enough to embrace the activities of the highest form
  of life; covers all those of the lower creatures。  The lowest
  plant; or animalcule; feeds; grows; and reproduces its kind。  In
  addition; all animals manifest those transitory changes of form
  which we class under irritability and contractility; and; it is
  more than probable; that when the vegetable world is thoroughly
  explored; we shall find all plants in possession of the same
  powers; at one time or other of their existence。
  I am not now alluding to such phaenomena; at once rare and
  conspicuous; as those exhibited by the leaflets of the sensitive
  plants; or the stamens of the barberry; but to much more widely
  spread; and at the same time; more subtle and hidden;
  manifestations of vegetable contractility。  You are doubtless aware
  that the common nettle owes its stinging property to the
  innumerable stiff and needle…like; though exquisitely delicate;
  hairs which cover its surface。  Each stinging…needle tapers from a
  broad base to a slender summit; which; though rounded at the end;
  is of such microscopic fineness that it readily penetrates; and
  breaks off in; the skin。  The whole hair consists of a very
  delicate outer case of wood; closely applied to the inner surface
  of which is a layer of semi…fluid matter; full of innumerable
  granules of extreme minuteness。  This semi…fluid lining is
  protoplasm; which thus constitutes a kind of bag; full of a limpid
  liquid; and roughly corresponding in form with the interior of the
  hair which it fills。  When viewed with a sufficiently high
  magnifying power; the protoplasmic layer of the nettle hair is seen
  to be in a condition of unceasing activity。  Local contractions of
  the whole thickness of its substance pass slowly and gradually from
  point to point; and give rise to the appearance of progressive
  waves; just as the bending of successive stalks of corn by a breeze
  produces the apparent billows of a cornfield。
  But; in addition to these movements; and independently of them; the
  granules are driven; in relatively rapid streams; through channels
  in the protoplasm which seem to have a considerable amount of
  persistence。  Most commonly; the currents in adjacent parts of the
  protoplasm take similar directions; and; thus; there is a general
  stream up one side of the hair and down the other。  But this does
  not prevent the existence of partial currents which take different
  routes; and sometimes trains of granules may be seen coursing
  swiftly in opposite directions within a twenty…thousandth of an
  inch of one another; while; occasionally; opposite streams come
  into direct collision; and; after a longer or shorter struggle; one
  predominates。  The cause of these currents seems to lie in
  contractions of the protoplasm which bounds the channels in which
  they flow; but which are so minute that the best microscopes show
  only their effects; and not themselves。
  The spectacle afforded by the wonderful energies prisoned within
  the compass of the microscopic hair of a plant; which we commonly
  regard as a merely passive organism; is not easily forgotten by one
  who has watched its display; continued hour after hour; without
  pause or sign of weakening。  The possible complexity of many other
  organic forms; seemingly as simple as the protoplasm of the nettle;
  dawns upon one; and the comparison of such a protoplasm to a body
  with an internal circulation; which has been put forward by an
  eminent physiologist; loses much of its startling character。
  Currents similar to those of the hairs of the nettle have been
  observed in a great multitude of very different plants; and weighty
  authorities have suggested that they probably occur; in more or
  less perfection; in all young vegetable cells。  If such be the
  case; the wonderful noonday silence of a tropical forest is; after
  all; due only to the dulness of our hearing; and could our ears
  catch the murmur of these tiny Maelstroms; '96' as they whirl in the
  innumerable myriads of living cells which constitute each tree; we
  should be stunned; as with the roar of a great city。
  Among the lower plants; it is the rule rather than the exception;
  that contractility should be still more openly manifested at some
  periods of their existence。  The protoplasm of Algae and Fungi
  becomes; under many circumstances; partially; or completely; freed
  from its woody case; and exhibits movements of its whole mass; or
  is propelled by the contractility of one; or more; hair…like
  prolongations of its body; which are called vibratile cilia。  And;
  so far as the conditions of the manifestation of the phaenomena of
  contractility have yet been studied; they are the same for the
  plant as for the animal。  Heat and electric shocks influence both;
  and in the same way; though it may be in different degrees。  It is
  by no means my intention to suggest that there is no difference in
  faculty between the lowest plant and the highest; or between plants
  and animals。  But the difference between the powers of the lowest
  plant; or animal; and those of the highest; is one of degree; not
  of kind; and depends; as Milne…Edwards '97' long ago so well pointed
  out; upon the extent to which the principle of the division of
  labour is carried out in the living economy。  In the lowest
  organism all parts are competent to perform all functions; and one
  and the same portion of protoplasm may successfully take on the
  function of feeding; moving; or reproducing apparatus。  In the
  highest; on the contrary; a great number of parts combine to
  perform each function; each part doing its allotted share of the
  work with great accuracy and efficiency; but being useless for any
  other purpose。
  On the other hand; notwithstanding all the fundamental resemblances
  which exist between the powers of the protoplasm in plants and in
  animals; they present a striking difference (to which I shall
  advert more at length presently); in the fact that plants can
  manufacture fresh protoplasm out of mineral compounds; whereas
  animals are obliged to procure it ready made; and hence; in the
  long run; depend upon plants。  Upon what condition this difference
  in the powers of the two great divisions of the world of life
  depends; nothing is at present known。
  With such qualifications as arises '98' out of the last…mentioned
  fact; it may be truly said that the acts of all living things are
  fundamentally one。  Is any such unity predicable of their forms?
  Let us seek in easily verified facts for a reply to this question。
  If a drop of blood be drawn by pricking one's finger; and viewed
  with proper precautions; and under a sufficiently high microscopic
  power; there will be seen; among the innumerable multitude of
  little; circular; discoidal bodies; or corpuscles; which float in
  it and give it its colour; a comparatively small number of
  colourless corpuscles; of somewhat larger size and very irregular
  shape。  If the drop of blood be kept at the temperature of the
  body; these colourless corpuscles will be seen to exhibit a
  marvellous activity; changing their forms with great rapidity;
  drawing in and thrusting out prolongations of their substance; and
  creeping about as if they were independent organisms。
  The substance which is thus active is a mass of protoplasm; and its
  activity differs in detail; rather than in principle; from that of
  the protoplasm of the nettle。  Under sundry circumstances the
  corpuscle dies and becomes distended into a round mass; in the
  midst of which is seen a smaller spherical body; which existed; but
  was more or less hidden; in the living corpuscle; and is called its
  nucleus。  Corpuscles of essentially similar structure are to be
  found in the skin; in the lining of the mouth; and scattered
  through the whole framework of the body。  Nay; more; in the
  earliest condition of the human organism; in that state in which it
  has but just become distinguishable from the egg in which it
  arises; it is nothing but an aggregation of such corpuscles; and
  every organ of the body was; once; no more than such an
  aggregation。
  Thus a nucleated mass of protoplasm tur