limits　of　species；　and；　what　is　more；　that　when　a　variety　has　come　into
  existence　in　nature；　there　are　natural　causes　and　conditions；　which　are
  amply　competent　to　play　the　part　of　a　selective　breeder；　and　although
  that　is　not　quite　the　evidence　that　one　would　like　to　havethough　it
  is　not　direct　testimonyyet　it　is　exceeding　good　and　exceedingly
  powerful　evidence　in　its　way。
  As　to　the　first　point；　of　varieties　existing　among　natural　species；　I
  might　appeal　to　the　universal　experience　of　every　naturalist；　and　of
  any　person　who　has　ever　turned　any　attention　at　all　to　the
  characteristics　of　plants　and　animals　in　a　state　of　nature；　but　I　may　as
  well　take　a　few　definite　cases；　and　I　will　begin　with　Man　himself。
  I　am　one　of　those　who　believe　that；　at　present；　there　is　no　evidence
  whatever　for　saying；　that　mankind　sprang　originally　from　any　more　than
  a　single　pair；　I　must　say；　that　I　cannot　see　any　good　ground　whatever；
  or　even　any　tenable　sort　of　evidence；　for　believing　that　there　is　more
  than　one　species　of　Man。　　Nevertheless；　as　you　know；　just　as　there　are
  numbers　of　varieties　in　animals；　so　there　are　remarkable　varieties　of
  men。　　I　speak　not　merely　of　those　broad　and　distinct　variations　which
  you　see　at　a　glance。　　Everybody；　of　course；　knows　the　difference
  between　a　Negro　and　a　white　man；　and　can　tell　a　Chinaman　from　an
  Englishman。　　They　each　have　peculiar　characteristics　of　colour　and
  physiognomy；　but　you　must　recollect　that　the　characters　of　these　races
  go　very　far　deeperthey　extend　to　the　bony　structure；　and　to　the
  characters　of　that　most　important　of　all　organs　to　usthe　brain；　so
  that；　among　men　belonging　to　different　races；　or　even　within　the　same
  race；　one　man　shall　have　a　brain　a　third；　or　half；　or　even　seventy　per
  cent。　bigger　than　another；　and　if　you　take　the　whole　range　of　human
  brains；　you　will　find　a　variation　in　some　cases　of　a　hundred　per　cent。
  Apart　from　these　variations　in　the　size　of　the　brain；　the　characters　of
  the　skull　vary。　　Thus　if　I　draw　the　figures　of　a　Mongul　and　of　a　Negro
  head　on　the　blackboard；　in　the　case　of　the　last　the　breadth　would　be
  about　seven…tenths；　and　in　the　other　it　would　be　nine…tenths　of　the
  total　length。　　So　that　you　see　there　is　abundant　evidence　of　variation
  among　men　in　their　natural　condition。　　And　if　you　turn　to　other　animals
  there　is　just　the　same　thing。　　The　fox；　for　example；　which　has　a　very
  large　geographical　distribution　all　over　Europe；　and　parts　of　Asia；　and
  on　the　American　Continent；　varies　greatly。　　There　are　mostly　large
  foxes　in　the　North；　and　smaller　ones　in　the　South。　　In　Germany　alone；
  the　foresters　reckon　some　eight　different　sorts。
  Of　the　tiger；　no　one　supposes　that　there　is　more　than　one　species；　they
  extend　from　the　hottest　parts　of　Bengal；　into　the　dry；　cold；　bitter
  steppes　of　Siberia；　into　a　latitude　of　50　degrees；so　that　they　may
  even　prey　upon　the　reindeer。　　These　tigers　have　exceedingly　different
  characteristics；　but　still　they　all　keep　their　general　features；　so　that
  there　is　no　doubt　as　to　their　being　tigers。　　The　Siberian　tiger　has　a
  thick　fur；　a　small　mane；　and　a　longitudinal　stripe　down　the　back；　while
  the　tigers　of　Java　and　Sumatra　differ　in　many　important　respects　from
  the　tigers　of　Northern　Asia。　　So　lions　vary；　so　birds　vary；　and　so；　if
  you　go　further　back　and　lower　down　in　creation；　you　find　that　fishes
  vary。　　In　different　streams；　in　the　same　country　even；　you　will　find
  the　trout　to　be　quite　different　to　each　other　and　easily　recognisable　by
  those　who　fish　in　the　particular　streams。　　There　is　the　same
  differences　in　leeches；　leech　collectors　can　easily　point　out　to　you
  the　differences　and　the　peculiarities　which　you　yourself　would　probably
  pass　by；　so　with　fresh…water　mussels；　so；　in　fact；　with　every　animal
  you　can　mention。
  In　plants　there　is　the　same　kind　of　variation。　　Take　such　a　case　even　as
  the　common　bramble。　The　botanists　are　all　at　war　about　it；　some　of　them
  wanting　to　make　out　that　there　are　many　species　of　it；　and　others
  maintaining　that　they　are　but　many　varieties　of　one　species；　and　they
  cannot　settle　to　this　day　which　is　a　species　and　which　is　a　variety！
  So　that　there　can　be　no　doubt　whatsoever　that　any　plant　and　any　animal
  may　vary　in　nature；　that　varieties　may　arise　in　the　way　I　have
  described；as　spontaneous　varieties；and　that　those　varieties　may　be
  perpetuated　in　the　same　way　that　I　have　shown　you　spontaneous　varieties
  are　perpetuated；　I　say；　therefore；　that　there　can　be　no　doubt　as　to　the
  origin　and　perpetuation　of　varieties　in　nature。
  But　the　question　now　is：Does　selection　take　place　in　nature？　　is　there
  anything　like　the　operation　of　man　in　exercising　selective　breeding；
  taking　place　in　nature？　　You　will　observe　that；　at　present；　I　say
  nothing　about　species；　I　wish　to　confine　myself　to　the　consideration　of
  the　production　of　those　natural　races　which　everybody　admits　to　exist。
  The　question　is；　whether　in　nature　there　are　causes　competent　to
  produce　races；　just　in　the　same　way　as　man　is　able　to　produce　by
  selection；　such　races　of　animals　as　we　have　already　noticed。
  When　a　variety　has　arisen；　the　CONDITIONS　OF　EXISTENCE　are　such　as　to
  exercise　an　influence　which　is　exactly　comparable　to　that　of　artificial
  selection。　　By　Conditions　of　Existence　I　mean　two　things；there　are
  conditions　which　are　furnished　by　the　physical；　the　inorganic　world；
  and　there　are　conditions　of　existence　which　are　furnished　by　the
  organic　world。　　There　is；　in　the　first　place；　CLIMATE；　under　that　head
  I　include　only　temperature　and　the　varied　amount　of　moisture　of
  particular　places。　　In　the　next　place　there　is　what　is　technically
  called　STATION；　which　meansgiven　the　climate；　the　particular　kind　of
  place　in　which　an　animal　or　a　plant　lives　or　grows；　for　example；　the
  station　of　a　fish　is　in　the　water；　of　a　fresh…water　fish　in　fresh
  water；　the　station　of　a　marine　fish　is　in　the　sea；　and　a　marine　animal
  may　have　a　station　higher　or　deeper。　　So　again　with　land　animals：　the
  differences　in　their　stations　are　those　of　different　soils　and
  neighbourhoods；　some　being　best　adapted　to　a　calcareous；　and　others　to
  an　arenaceous　soil。　　The　third　condition　of　existence　is　FOOD；　by　which
  I　mean　food　in　the　broadest　sense；　the　supply　of　the　materials　necessary
  to　the　existence　of　an　organic　being；　in　the　case　of　a　plant　the
  inorganic　matters；　such　as　carbonic　acid；　water；　ammonia；　and　the
  earthy　salts　or　salines；　in　the　case　of　the　animal　the　inorganic　and
  organic　matters；　which　we　have　seen　they　require；　then　these　are　all；
  at　least　the　two　first；　what　we　may　call　the　inorganic　or　physical
  conditions　of　existence。　　Food　takes　a　mid…place；　and　then　come　the
  organic　conditions；　by　which　I　mean　the　conditions　which　depend　upon　the
  state　of　the　rest　of　the　organic　creation；　upon　the　number　and　kind　of
  living　beings；　with　which　an　animal　is　surrounded。　　You　may　class　these
  under　two　heads：　there　are　organic　beings；　which　operate　as
  'opponents'；　and　there　are　organic　beings　which　operate　as　'helpers'　to
  any　given　organic　creature。　　The　opponents　may　be　of　two　kinds：　there
  are　the　'indirect　opponents'；　which　are　what　we　may　call　'rivals'；　and
  there　are　the　'direct　opponents'；　those　which　strive　to　destroy　the
  creature；　and　these　we　call　'enemies'。　By　rivals　I　mean；　of　course；　in
  the　case　of　plants；　those　which　require　for　their　support　the　same　kind
  of　soil　and　station；　and；　among　animals；　those　which　require　the　same
  kind　of　station；　or　food；　or　climate；　those　are　the　indirect　opponents；
  the　direct　opponents　are；　of　course；　those　which　prey　upon　an　animal　or
  vegetable。　　The　'helpers'　may　also　be　regarded　as　direct　and　indirect：
  in　the　case　of　a　carnivorous　animal；　for　example；　a　particular
  herbaceous　plant　may　in　multiplying　be　an　indirect　helper；　by　enabling
  the　herbivora　on　which　the　carnivore　preys　to　get　more　food；　and　thus
  to　nourish　the　carnivore　more　abundantly；　the　direct　helper　may　be　best
  illustrated　by　reference　to　some　parasitic　creature；　such　as　the
  tape…worm。　　The　tape…worm　exists　in　the　human　intestines；　so　that　the
  fewer　there　are　of　men　the　fewer　there　will　be　of　tape…worms；　other
  things　being　alike。　　It　is　a　humiliating　reflection；　perhaps；　that　we
  may　be　classed　as　direct　helpers　to　the　tape…worm；　but　the　fact　is　so：
  we　can　all　see　that　if　there　were　no　men　there　would　be　no　tape…worms。
  It　is　extremely　difficult　to　estimate；　in　a　proper　way；　the　importance
  and　the　working　of　the　Conditions　of　Existence。　　I　do　not　think　there
  were　any　of　us　who　had　the　remotest　notion　of　properly　estimating　them
  until　the　publication　of　Mr。　Darwin's　work；　which　has　placed　them
  before　us　with　remarkable　clearness；　and　I　must　endeavour；　as　far　as　I
  can　in　my　own　fashion；　to　give　you　some　notion　of　how　they　work。　　We
  shall　find　it　easiest　to　take　a　simple　case；　and　one　as　free　as
  possible　from　every　kind　of　complication。
  I　will　suppose；　therefore；　that　all　the　habitable　part　of　this
  globethe　dry　land；　amounting　to　about　51；000；000　square　miles；I
  will　suppose　that　the　whole　of　that　dry　land　has　the　same　climate；　and
  that　it　is　composed　of　the　same　kind　of　rock　or　soil；　so　that　there　will
  be　the　same　station　everywhere；　we　thus　get　rid　of　the　peculiar
  in